Calculate evenness over a gridded map or as a time series (see 'Details' for more information).
Usage
pielou_evenness_map(data, ...)
pielou_evenness_ts(data, ...)
williams_evenness_map(data, ...)
williams_evenness_ts(data, ...)
Arguments
- data
A data cube object (class 'processed_cube').
- ...
Arguments passed on to
compute_indicator_workflow
ci_type
(Optional) Type of bootstrap confidence intervals to calculate. (Default: "norm"). Select "none" to avoid calculating bootstrap CIs.
cell_size
(Optional) Length of grid cell sides, in km or degrees. If NULL, this will be automatically determined according to the geographical level selected. This is 100 km or 1 degree for 'continent' or 'world', 10 km or (for a degree-based CRS) the native resolution of the cube for 'country', 'sovereignty' or 'geounit'. If level is set to 'cube', cell size will be the native resolution of the cube for a degree-based CRS, or for a km-based CRS, the cell size will be determined by the area of the cube: 100 km for cubes larger than 1 million sq km, 10 km for cubes between 10 thousand and 1 million sq km, 1 km for cubes between 100 and 10 thousand sq km, and 0.1 km for cubes smaller than 100 sq km. (Default: NULL)
level
(Optional) Spatial level: 'cube', 'continent', 'country', 'world', 'sovereignty', or 'geounit'. (Default: 'cube')
region
(Optional) The region of interest (e.g., "Europe"). This parameter is ignored if level is set to 'cube' or 'world'. (Default: NULL)
ne_type
(Optional) The type of Natural Earth data to download: 'countries', 'map_units', 'sovereignty', or 'tiny_countries'. This parameter is ignored if level is set to 'cube' or 'world'. (Default: "countries")
ne_scale
(Optional) The scale of Natural Earth data to download: 'small' - 110m, 'medium' - 50m, or 'large' - 10m. (Default: "medium")
output_crs
(Optional) The CRS you want for your calculated indicator. (Leave blank to let the function choose a default based on grid reference system.)
first_year
(Optional) Exclude data before this year. (Uses all data in the cube by default.)
last_year
(Optional) Exclude data after this year. (Uses all data in the cube by default.)
spherical_geometry
(Optional) If set to FALSE, will temporarily disable spherical geometry while the function runs. Should only be used to solve specific issues. (Default is TRUE).
make_valid
(Optional) Calls st_make_valid() from the sf package after creating the grid. Increases processing time but may help if you are getting polygon errors. (Default is FALSE).
num_bootstrap
(Optional) Set the number of bootstraps to calculate for generating confidence intervals. (Default: 100)
shapefile_path
(optional) Path of an external shapefile to merge into the workflow. For example, if you want to calculate your indicator particular features such as protected areas or wetlands.
shapefile_crs
(Optional) CRS of a .wkt shapefile. If your shapefile is .wkt and you do NOT use this parameter, the CRS will be assumed to be EPSG:4326 and the coordinates will be read in as lat/long. If your shape is NOT a .wkt the CRS will be determined automatically.
invert
(optional) Calculate an indicator over the inverse of the shapefile (e.g. if you have a protected areas shapefile this would calculate an indicator over all non protected areas within your cube). Default is FALSE.
include_land
(Optional) Include occurrences which fall within the land area. Default is TRUE. *Note that this purely a geographic filter, and does not filter based on whether the occurrence is actually terrestrial. Grid cells which fall partially on land and partially on ocean will be included even if include_land is FALSE. To exclude terrestrial and/or freshwater taxa, you must manually filter your data cube before calculating your indicator.
include_ocean
(Optional) Include occurrences which fall outside the land area. Default is TRUE. Set as "buffered_coast" to include a set buffer size around the land area rather than the entire ocean area. *Note that this is purely a geographic filter, and does not filter based on whether the occurrence is actually marine. Grid cells which fall partially on land and partially on ocean will be included even if include_ocean is FALSE. To exclude marine taxa, you must manually filter your data cube before calculating your indicator.
buffer_dist_km
(Optional) The distance to buffer around the land if include_ocean is set to "buffered_coast". Default is 50 km.
force_grid
(Optional) Forces the calculation of a grid even if this would not normally be part of the pipeline, e.g. for time series. This setting is required for the calculation of rarity, and is turned on by the ab_rarity_ts and area_rarity_ts wrappers. (Default: FALSE)
Value
An S3 object with the classes 'indicator_map' or 'indicator_ts' and 'pielou_evenness' or 'williams_evenness' containing the calculated indicator values and metadata.
Details
Evenness
Species evenness is a commonly used indicator that measures how uniformly individuals are distributed across species in a region or over time. It provides a complement to richness by taking relative abundance into account. Although GBIF provides information about abundances as individual counts, the majority of entries lack this information. Hence, evenness can only be calculated using the proportions of observations rather than proportions of individuals. Strictly speaking, the evenness measures therefore indicate how uniformly species are represented in the respective data set rather than the true evenness of the ecological community.
Pielou's evenness
Pielou's evenness (1966) is a well-known and commonly used evenness measure. It is calculated as:
$$ E = \frac{-\sum_{i=1}^{S} p_i \ln(p_i)}{\ln(S)} $$ where S is the number of species and pi is the proportion of occurrences represented by species i.
Williams' evenness
An analysis of evenness properties by Kvålseth (2015) showed that an evenness index introduced by Williams in 1977 in an unpublished manuscript has two important properties which Pielou's does not. The properties in question are complex mathematical properties known as the Schur-Concavity and value validity, but we attempt to describe them here more simply. If a measure of evenness is Schur-concave, it means that when the distribution of individuals becomes more evenly spread across species, the measure of evenness will stay the same or increase, but never decrease. Value validity means that an evenness index should provide sensible and meaningful values across its range for any given distribution of species abundances. Kvålseth referred to this evenness measure as E9 but we refer to it as Williams' evenness.
Williams' evenness is calculated as:
$$ 1 - \sqrt{\frac{S\sum_{i=1}^{S} p_i^2 - 1}{S - 1}} $$
where S is the number of species and pi is the proportion of occurrences represented by species i.
Functions
pielou_evenness_map()
:pielou_evenness_ts()
:williams_evenness_map()
:williams_evenness_ts()
:
References
Pielou, E. C. (1966). The measurement of diversity in different types of biological collections. Journal of theoretical biology, 13, 131-144.
Kvålseth, T. O. (2015). Evenness indices once again: critical analysis of properties. SpringerPlus, 4, 1-12.
Examples
if (FALSE) { # \dontrun{
pe_map <- pielou_evenness_map(example_cube_1, level = "country",
region = "Denmark")
plot(pe_map)
} # }
if (FALSE) { # \dontrun{
pe_ts <- pielou_evenness_ts(example_cube_1, first_year = 1985)
plot(pe_ts)
} # }
if (FALSE) { # \dontrun{
we_map <- williams_evenness_map(example_cube_1, level = "country",
region = "Denmark")
plot(we_map)
} # }
if (FALSE) { # \dontrun{
we_ts <- williams_evenness_ts(example_cube_1, first_year = 1985)
plot(we_ts)
} # }